Initial values for parameters: for nuclear genes, for cpDNA, for mtDNA or as follows: initial values of nucleotide exchangeabilities [m_ij]: ratio (m) of double/triple to single/double nucleotide changes: _0.0__ m_tc|ag /m_[tc][ag]: _1.0__ m_ag / m_tc[ag]: _1.0__ m_ta / m_[tc][ag]: _1.0__ m_tg / m_[tc][ag]: _1.0__ m_ca / m_[tc][ag]: _1.0__ initial values of nucleotide equilibrium frequencies [f_i]: f_a: 0.25 f_c: 0.25 f_g: 0.25